小智和皮卡丘等一行伙伴在旅途中前行。某日,宝可梦波尔凯尼恩从天而降并被迫与小智锁在了一起。波尔凯尼恩与被人类伤害过的宝可梦们共同生活,因此非常讨厌人类,但这条神奇的锁链却让他们无法分开,奇妙的故事就此开始。小智与皮卡丘又将一起开始新的冒险…… 另一方面,在超级机关城市阿佐特王国,大臣贾维斯得到了常年追求的终极发明——500 年前诞生于阿佐特王国的人造宝可梦玛机雅娜。据说在玛机雅娜体内,有被称为 “魂心”的未知力量,贪婪又邪恶的贾维斯企图利用这股力量来统治王国。小智和波尔凯尼恩将会如何化解彼此的嫌隙?波尔凯尼恩与玛机雅娜又有着怎样的渊源?他们要如何共同对抗贾维斯拯救阿佐特王国?玛机雅娜的最终命运又将如何结局?一场精灵宝可梦史上最伟大的战役即将展开!
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